docOrder Family Genus Species Author Year Exx. Region Coleoptera
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Order Family Genus Species Author Year Exx. Region Coleoptera
Order Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Family Ceratocanthidae Ceratocanthidae Ceratocanthidae Ceratocanthidae Ceratocanthidae Ceratocanthidae Diphyllostomatidae Diphyllostomatidae Diphyllostomatidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Genus Germarostes Germarostes Germarostes Germarostes Germarostes Haroldostes Diphyllostoma Diphyllostoma Diphyllostoma Anoplotrupes Anoplotrupes Anoplotrupes Athyreus Athyreus Blackburnium Blackburnium Bolbelasmmus Bolbocerastes Bolbocerastes Bolbocerastes Bolbocerosoma Bolbocerosoma Bolbocerosoma Bolbocerosoma Bolbocerosoma Bolbocerosoma Bolbocerosoma Bolbochromus Bolborhombus Bradycinetulus Bradycinetulus Ceratophyus Cnemotrupes Cnemotrupes Cnemotrupes Enoplotrupes Eogeotrupes Epigeotrupes Eucanthus Eucanthus Eucanthus Eucanthus Eucanthus Frickius Geohowdenius Geohowdenius Geotrupes Geotrupes Geotrupes Glyptogeotrupes Lethrus Lethrus Lethrus Lethrus Lethrus Lethrus Lethrus Lethrus Lethrus Mycotrupes Mycotrupes Neoathyreus Species aphodioides globosus macleayi plicatus punctulatus senegalensis fimbriata linsleyi nigricollis ballyi hornii stercorosus excavatus sp. insigne sloanei horni imperialis regalis serratus biplagiatum bruneri confusum farctum hamatum pusillum tumefactum n.sp. sallaei ferrugineus fossatus polyceros blackburnii semiopacus splendidus sharpi auratus laevistriatus alutaceus greeni impressus lazarus lazarus variolosus egeriei opacus mutator spiniger stercorarius impressus acutangulus apterus apterus cephalotus inermis karelini koshantschikovi scoparius sulcipennis gaigei retusus excavatus Author Illiger Say Perty Germar Ohaus Castelnau Fall Fall Fall Jekel Blanchard Scriba Castelnau Lea Blackburn Rivers Cartwright Cartwright LeConte Dawson and McCollock Dawson and McCollock Brown Fabricius Brown Dawson and McCollock Beauvois Bates Beauvois Haldeman Pallas Fabricius Jekel Fabricius Jordan and Rothschild Motschulsky Motschulsky Cartwright Robinson Howden Fabricius Germain Germar Haldeman Marsham Marsham Linnaeus Gebler Ballion Laxm. Fischer Pallas Reitter Gebler Semenov Fischer Kraatz Olsen and Hubbel LeConte Laporte Year Exx. Region 1800 71 nea 1835 33 nea 1830 1 pal 1843 3 pal 1911 1 pal 1840 1 pal 1901 3 nea 1932 6 nea 1912 34 nea 1865 45 nea 1888 13 nea 1791 77 pal 1840 1 pal 1 pal 1916 2 aus 1889 1 aus 1886 1 nea 1953 9 nea 1953 1 nea 1854 10 nea 1924 4 nea 1924 8 nea 1928 2 nea 1775 9 nea 1929 3 1924 1 nea 1805 8 nea 1 ori 1887 9 nea 1805 1 nea 1853 5 nea 1771 2 pal 1781 28 nea 1865 45 nea 1781 40 nea 1893 2 pal 1857 2 1 pal 1944 1 nea 6 nea 1964 10 nea 1775 109 nea 33 nea 1897 6 pal 1824 14 nea 1853 7 nea 1802 12 pal 1802 6 pal 1758 12 pal 1841 1 pal 1871 2 pal 1770 2 1822 2 pal 1771 4 neo 1897 1 neo 1845 2 neo 1892 1 neo 1820 2 neo 1883 2 neo 1954 1 nea 1866 1 nea 1840 1 pal Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Geotrupidae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaphyridae Glaresidae Glaresidae Glaresidae Glaresidae Hybosoridae Hybosoridae Hybosoridae Lucanidae Neoathyreus Neoathyreus Odonteus Odonteus Odonteus Odonteus Odonteus Odonteus Odonteus Odonteus Odonteus Odonteus Odonteus Odonteus Odonteus Odonteus Odonteus Odonteus Peltotrupes Sericotrupes Taurocerastes Thorectes Thorectes Thorectes Thorectes Trypocopris Trypocopris Trypocopris Trypocopris Typhoeus Typhoeus Amphicoma Amphicoma Amphicoma Amphicoma Amphicoma Amphicoma Amphicoma Amphicoma Amphicoma Amphicoma Amphicoma Amphicoma Amphicoma Amphicoma Amphicoma Cratoscelis Cratoscelis Glaphyrus Glaphyrus Glaphyrus Lichanthe Lichanthe Lichanthe Lichnia Glaresis Glaresis Glaresis Glaresis Coelodes Hybosorus Hybosorus Aegus fissicornis lanei binasutum capense corniger darlingtoni distinguendum falli floridanus gallicus laesicolle liebecki obesus simi sp. sp. striatopunctatum thoracicornis profundus niger patagonicus geminatus intermedius marginatus rugatulus pyrenaeus vernalis vernalis vernalis typhoeus typhoeus acrtos arctos bombyliformis bombylius fastuosa hyrax kuschakewitschi lasserrei malatia papaveris vittata vittata vittatta vulpes vulpes villosa vulpina festivus maurus micans rathvoni ursina vulpina limbata ecostata gordoni phoenicia zvirgzdinsi ovalis illigeri spp. minutus Harold Martinez Fairmaire Klug Melsheimer Wallis Fairmaire Wallis Wallis Mulsant Fairmaire Wallis LeConte Wallis 1880 1952 1861 1843 1844 1928 1861 1928 1928 1842 1856 1928 1859 1928 Castelnau Wallis Howden Marsham Phillipi Gene Costa Poiret Jekel Charpentier Linnaeus Heer Kuster Linn. Linnaeus Faldermann Reitter Pallas Fabricius Reitter Truqui Ball Germar Truqui Sturm Fabricius Reitter Reitter Brulle Fabricius Blanchard Erichson Menier Linnaeus Faldermann LeConte LeConte Hentz Erichson Fall Warner Fall Warner Robinson Reiche 1840 1928 1952 1802 1866 1839 1827 1787 1865 1825 1758 1847 1853 1758 1758 1835 1890 1781 1789 1890 1848 1871 1834 1848 1843 1775 1890 1890 Gestro 1792 1850 1835 1836 1758 1835 1863 1861 1826 1851 1907 1907 1853 1 2 2 2 4 4 1 4 31 3 1 34 20 3 1 1 1 2 5 2 2 3 9 2 2 2 13 1 4 8 8 1 1 1 8 1 1 1 17 7 1 1 2 1 1 2 7 2 1 3 1 13 4 16 4 1 2 1 2 pal pal pal afr nea nea pal nea nea neo pal nea nea nea nea ori nea nea pal pal pal pal pal pal pal pal pal pal pal pal pal pal pal pal pal pal pal pal pal pal pal neo neo neo pal neo neo pal afr pal nea nea nea neo nea nea nea nea 8 nea 18 1 Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Aesalus Apterocyclus Apterodorcus Bomansodus Brasilucanus Casignetus Ceratognathus Ceruchus Ceruchus Ceruchus Ceruchus Chalcodes Chiasognathinus Chiasognathus Cladognathus Cladognathus Cladognathus Cyclommatus Cyclommatus Dorcus Dorcus Dorcus Dorcus Dorcus Eurytrachellelus Eurytrachellelus Eurytrachellelus Eurytrachellelus Eurytrachellelus Eurytrachellelus Eurytrachellelus Eurytrachellelus Eurytrachellelus Eurytrachellelus Eurytrachellelus Eurytrachellelus Eurytrachellelus Eurytrachellelus Figulus Gnaphaloryx Hemisodorcus Hexarthrius Hexarthrius Lamprima Lamprima Lamprima Lamprima Leptinopterus Leptinopterus Lissapterus Lissotes Lissotes Lucanus Lucanus Lucanus Metopodontus Metopodontus Metopodontus Metopodontus Metopodontus Metopodontus Neolucanus Neolucanus scarabaeoides honoluluensis bacchus impubis alvaregei humboldti curvidens chrysomelinus piceus punctatus striatus carinatus gaujoni granti confucius confucius giraffa frey-gessneri tarandus brevis immundus parallelepipedus parallelepipedus parallelus alcides arfakianus bucephalus cribriceps egregius gypaetus intermedius reichei rubrofemoratus saiga striatipennis submolaris titanus tityus foveicollis opacus napalensis deyrollei parryi adolphinae aurata latreille varians ibex tibialis howittanus reticulatus spp. cervus elaphus smithi bison cinctus foveatus occipitalis savagei spp. alces baladeva Panzer Waterhouse Westwood Parry Vulcano and Pereira Gyllenhal Broun Hochenwarth Weber LeConte LeConte Linnaeus Oberthuer Stephens Hope Olivier Ritsema Thuenberg Say Arrow Linnaeus Ganglbauer Say Vollenhoven Lansberge Perty Chevrolat Moellenkamp Castelnau Gestro Hope Vollenhoven Olivier Motschulsky Westwood Boisduval Hope Boisduval Burmeister Hope Parry Hope Gestro Latreille McLeay Germar Billberg Eschscholtz Westwood Westwood Linnaeus Fabricius Parry Olivier Montrouzier Hope Westwood Hope Fabricius Hope 1794 1904 1785 1801 1869 1859 1 1 4 4 1 1 2 2 297 3 9 2 1 10 1 1 4 1 15 4 1 21 3 82 1 2 5 3 2 1 26 1 4 10 7 1 6 1 2 1 1 1 1 2 2 1 1 2 5 1 2 2 53 7 3 7 1 1 1 5 5 4 5 neo aus neo nea nea nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Lucanidae Ochodaeidae Ochodaeidae Ochodaeidae Ochodaeidae Ochodaeidae Ochodaeidae Ochodaeidae Neolucanus Nicagus Nigidius Nigidius Nigidius Nigidius Odontolabis Odontolabis Odontolabis Odontolabis Odontolabis Odontolabis Odontolabis Odontolabis Odontolabis Phalacrognathus Platyceroides Platyceroides Platyceroides Platyceropsis Platycerus Platycerus Platycerus Platycerus Platycerus Platycerus Platycerus Prismognathus Prismognathus Prosopocoelus Prosopocoelus Prosopocoelus Prosopocoelus Prosopocoelus Prosopocoelus Prosopocoelus Psalidoremus Pseudolucanus Pseudolucanus Pseudolucanus Pseudolucanus Pycnosiphorus Pycnosiphorus Pycnosiphorus Pycnosiphorus Pycnosiphorus Pycnosiphorus Sclerostomus Sclerostomus Sinodendron Sinodendron Sphenognathus Sphenognathus Streptocerus Streptocerus undetermined Ochodaeus Ochodaeus Ochodaeus Ochodaeus Ochodaeus Ochodaeus Ochodaeus castanopterus obscurus bonneuili laevicollis medimniennsis spp. bellicosus cuvera dalmani delesserti gazella moellenkampi siva sommeri wollastoni muelleri aeneus agassii laticollis keeni caraboides marginalis marginalis oregonensis piceus spp. virescens angularis subaeneus dorsalis hasterti lorquini natalensis senegalensis serricornis spp. inclinatus capreolus mazama placidus spp. caelatus costatus fasciatus lessoni mandibularis virgatus cucullatus tuberculatus cylindricum rugosum higginsi nobilis speciosus speciosus spp. biarmatus chrysomeloides frontalis gnatho inarmatus kansanus n.sp. nr.biarmatus Hope LeConte Boileau Westwood Seguy 1848 Castelnau HOpe Westwood Guerin Fabricius Fruhstorfer Westwood Parry Parry Kriesche VanDyke LeConte Casey Casey Linnaeus Casey Benesh Westwood Kirby Fabricius Waterhouse Motschulsky Erichson Boileau Deyrolle Waterhouse Klug Latreille Motschulsky Linnaeus LeConte Say Blanchard Benesh Germain Buquet Solier Nagel Blanchard Solier Linnaeus Mannerheim Parry Parry Fairaire Fairmaire LeConte Schrank LeConte Fall Schaeffer Fall 1758 1843 1868 1781 1863 1907 1906 1909 5 15 2 2 1 3 2 6 12 1 1 1 2 2 7 71 4 3 2 6 11 14 2 28 14 15 54 1 2 2 2 1 4 2 2 1 14 184 9 53 35 4 1 5 2 1 1 4 2 12 75 1 1 1 1 64 25 2 1 1 9 4 2 nea neo nea nea pal nea nea nea nea nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Ochodaeidae Ochodaeidae Ochodaeidae Ochodaeidae Ochodaeidae Ochodaeidae Ochodaeidae Ochodaeidae Ochodaeidae Pleocomidae Pleocomidae Pleocomidae Pleocomidae Pleocomidae Pleocomidae Pleocomidae Pleocomidae Pleocomidae Pleocomidae Pleocomidae Pleocomidae Pleocomidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Ochodaeus Ochodaeus Ochodaeus Ochodaeus Ochodaeus Ochodaeus Ochodaeus Ochodaeus Ochodaeus Pleocoma Pleocoma Pleocoma Pleocoma Pleocoma Pleocoma Pleocoma Pleocoma Pleocoma Pleocoma Pleocoma Pleocoma Pleocoma Actinobolus Adoretus Adoretus Adoretus Adoretus Adoretus Aegialia Aegialia Aegialia Aegialia Aegialia Aegialia Aegialia Aegialia Aegialia Aegialia Aegialia Aegialia Aegopsis Aegopsis Aethiessa Aethiessa Aethiessa Allomyrina Allorrhina Allorrhina Amaladera Amaurodes Amazula Amithao Amphimallon Amphimallon Amphimallon Amphimallon Amphimallon Amphimallon Amphimallon Amphimallon Amphimallon Amphimallon Anatropis n.sp. nr.biarmatus pectoralis peninsularis planifrons praesidii repandus simplex sparsus spp. australis badia behrensi crinita dubitalis edwardsi minor oregonensis puncticollis simi spp. staff tularensis radians compressus flaveolus nigrifrons sinicus tenuimaculatus arenaria arenaria blanchardi conferta convexa criddlei lacustris latispina opaca opifex punctata sabuleti curvicornis sp. floralis floralis floralis dichotomus carmelita scabriuscula euphorbiae passerini suavis decemguttatus assimilis ater ater caucasicus glabripennis majalis quercanus seidlitzi solstitialis solstitialis verticalis ? LeConte Horn Schaeffer Bates Fall LeConte LeConte 1868 1895 1906 1887 1909 1854 1868 Fall Fall LeConte Linsley Davis LeConte Linsley Leach Rivers Davis 1911 1917 1874 1938 1935 1874 1938 1933 1889 1934 Schaufuss Leach Westwood Weber Fahraeus Stephens Burmeister Waterhouse Fabricius Fabricius Horn Horn Fall Brown LeConte LeConte Brown Horn Brown Panzer Burmeister 1870 1933 1841 1801 1857 1809 1855 1875 1787 1787 1887 1871 1932 1931 1850 1878 1931 1887 1931 1797 1847 Fabricius Herbst Gory and Percheron Linnaeus Burmeister Swederus Burmeister Westwood Burmeister Waterhouse Herbst Herbst Olivier Gyllenhal Ballion Razoumowsky Burmeister Kraatz Linnaeus Gyllenhal Horn 1787 1790 1833 1771 1842 1787 1855 1844 1847 1876 1790 1790 1789 1817 1871 1789 1855 1891 1758 1817 1879 2 1 1 7 14 3 5 1 3 1 1 2 6 2 1 6 2 1 2 8 1 1 1 4 2 1 3 4 6 6 2 8 3 3 5 4 1 1 6 2 1 2 31 1 1 5 1 2 1 1 1 1 1 2 1 1 1 122 3 1 17 4 11 nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea neo afr afr afr ori ori nea pal nea nea nea nea nea nea nea nea nea pal neo neo afr pal afr ori neo neo pal afr afr neo pal pal pal pal pal nea pal pal pal pal nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Ancognatha Ancognatha Anisonyx Anisoplia Anisoplia Anisoplia Anisoplia Anisoplia Anisoplia Anisoplia Anisoplia Anisoplia Anisoplia Anisoplia Anisoplia Anisoplia Anisoplia Anisoplia Anisoplia Anochilia Anochilia Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala manca spp. hilaris agnata agricola agricola austriaca austriaca baetica bromicola farraria flavipennis flavipennis leucaspis remota segetum syriaca tempestiva tenebralis laevigata spp. arida atriplicis aulax binotata camancha cavifrons cinta cupricollis delicata digressa donovani dubia dubia dubia dubia dubia dubia flavilla flavilla flavipennis flavipennis flavipennis foraminosa foraminosa forreri fusciceps gemella histrionella hoepfneri immatura inconstans inconstans innuba lucicola ludoviciana marginata marginicollis maritima medorensis mendica minuta nigropicta LeConte 1866 Peringuey Reitter Poda Goeze Herbst Herbst Erichson Germar Erichson Brulle Erichson Laporte Reitter Herbst Burmeister Erichson Burmeister Gory and Percheron 1902 1889 1761 1777 1783 1783 1847 1817 1847 1832 1847 1840 1889 1783 1844 1847 1844 1835 Casey Fabricius Wiedemann Gyllenhal Wickham LeConte Bates Chevrolat Casey Casey Blanchard Scopoli Scopoli Scopoli Scopoli Fabricius Fabricius Bates LeConte Burmeister LeConte Casey Bates Bates Bates Fahraeus Say Bates Bates Bohemann Burmeister Burmeister Fabricius Fabricius Schaeffer Fabricius Bates Casey Casey Casey Burmeister Casey 1915 1787 1832 1817 1913 1868 1888 1915 1915 1850 1763 1763 1763 1763 1775 1775 1888 1863 1844 1854 1915 1888 1888 1888 1857 1835 1888 1888 1860 1844 1844 1787 1798 1906 1775 1888 1915 1915 1915 1844 1915 43 29 2 1 5 15 2 10 1 2 1 1 13 2 3 16 2 2 2 4 4 4 1 1 5 50 3 2 3 5 2 1 8 9 14 1 7 2 1 29 24 3 2 2 1 2 3 3 1 2 1 2 11 85 4 77 2 4 1 2 5 210 nea neo afr pal pal pal pal pal pal pal pal pal pal pal pal pal pal pal pal ori ori nea pal ori nea nea nea neo neo nea nea neo pal pal pal pal pal pal nea nea nea nea nea nea neo neo afr neo neo neo afr neo neo nea nea nea nea neo nea nea nea nea nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomala Anomiopsoides Anoplocheilus Anoplodrepanus Anoplodrepanus Anoplognathus Anoplognathus Anoplognathus Anoplognathus Anoplognathus Anoplognathus Anoplognathus Anoplognathus Anoplognathus Anoplosiagum Anoxia Anoxia Anoxia Anoxia Anoxia Anticheira Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius Aphodius oblivia orientalis orientalis osmanlis papagoana parvula pasadenae sagax semilivida semitonsa sp. nr.plurisulcata spp. spp. stigmotella tenera tenera undulata vanpatteni villosella vitis pereirai figuratus pecki reconditus macleayi montanus olivieri pallidicollis sp. sp. sp. suturalis viriditarsis sp. nr.rufum asiatica orientalis pilosa scutellaris villosa campisilvaticus aemulus albertanus aleutus aleutus aleutus alpinus alternatus alternatus angustus anthracinus aquilonarius ater badipes badipes bicolor bimaculatus bonvouloiri brasiliensis brimleyi browni bryanti campestris carri Horn Waterhouse Waterhouse Blanchard Casey Burmeister Casey Casey LeConte Bates Casey Casey Casey Melsheimer Bates Blanchard Fabricius Menetries Bohmann Howden Matthews Blackburn MacLeay Dalman Blanchard 1884 1875 1875 1850 1915 1844 1915 1878 1888 26 3 15 1 26 1 3 4 6 2 1915 1915 1915 1844 1888 1850 1775 6 11 3 3 21 45 4 1 10 1857 2 Boisduval Leach 1835 1815 2 5 5 1 8 2 1 3 3 1 Desbrocher Krynicki Fabricius Mulsant Fabricius Ohaus Horn Brown Eschscholtz Fall and Cockerell Horn Scopoli Horn Horn Hatch LeConte Brown Degeer Melsheimer Melshiemer Say Laxmann Harold Castelnau Cartwright Hinton Brown Blatchley Brown 1871 1832 1792 1842 1781 1905 1887 1928 1822 1907 1870 1763 1870 1879 1972 1878 1928 1774 1844 1844 1823 1770 1860 1840 1939 1934 1929 1912 1929 2 1 12 2 3 4 6 1 7 14 16 4 3 7 3 12 1 7 5 5 15 8 1 1 5 1 1 53 1 1892 1873 1817 1851 nea aus nea pal nea nea nea nea nea neo neo nea neo nea nea neo nea neo neo pal neo afr neo neo aus aus aus aus aus aus aus aus neo pal pal pal pal pal neo nea nea nea nea nea neo nea nea nea nea nea neo nea nea nea neo neo pal nea nea nea nea nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae 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constans contaminatus corruptor crassuloides crassulus cribrarius criddlei cuniculus denticulatus dentigerulus depressus depressus depressus distinctus distinctus duplex elegans erraticus erraticus erraticus erraticus erraticus explanatus femoralis fimetarius fimetarius fossor fucosus fuliginosus geomysi granarius granarius grombezewskyi guttatus haemorrhoidalis haemorrhoidalis haemorrhoidalis hamatus hydrochaeris hyperboreus ictericus immundus iowensis kirni lacustris larreae lentus lividus lugens lunifer luridus luridus luridus Brown Fall Panzer Horn Reitter Say Mannerheim LeConte Horn Creutzer Duftschmid Herbst Brown Fall and Cockerell Horn Brulle Brown Chevrolat Haldeman Brown Kugelann Felderman Erichson Muller Muller LeConte Allibert Linnaeus Mulsant Schmidt Mulsant Gebler LeConte Say Linnaeus Reitter Linnaeus Schmidt Harold Cartwright Linnaeus Mulsant Koshant Eschscholtz Linnaeus Linnaeus Linneaus Say Fabricius LeConte Laicharting Creutzer Wickham Cartwright LeConte Horn Horn Olivier Creutzer Solsky Fabricius Fabricius Muller 1927 1932 1798 1870 1892 1823 1853 1850 1887 1799 1805 1783 1929 1907 1870 1836 1928 1864 1848 1929 1792 1835 1848 1776 1776 1878 1847 1758 1842 1916 1842 1809 1878 1823 1758 1892 1758 1922 1863 1939 1767 1842 1891 1823 1758 1758 1758 1823 1798 1850 1781 1799 1913 1944 1850 1887 1870 1789 1799 1876 1775 1775 1776 4 9 6 106 7 7 55 10 4 6 5 1 2 15 1 5 1 9 2 4 1 6 11 6 169 1 1 118 3 5 2 8 1 4 469 5 121 13 1 1 401 1 nea nea neo nea neo nea nea nea nea neo neo neo nea nea nea neo nea nea nea nea neo neo neo neo nea nea neo nea neo neo neo neo nea nea nea neo nea nea pal nea nea neo neo nea neo nea neo nea neo nea neo neo nea nea nea nea nea nea neo neo 2 1 28 1 23 2 1 12 7 2 6 5 4 1 422 12 1 4 11 neo 5 neo Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae 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omissus omissus opacus opifex pardalis paykulli pectoralis peculiosus phaeopterus phaleroides piceatus piceus pictus pingellus pinguis prodromus prodromus pseudabusus pubescens punctissimus pusillus quadriguttatus quadriguttatus quadrimaculatus rectus rectus rossi rubeolus rubripennis rudis rudis ruficlaris ruficlarus rufipes rugatus rusicola sallei satellitus scrofa scrutator scuticollis scybalarius sepultus serval sigmoides snowi socialis sordidus sparsus Horn Haldeman Horn Schm. Schmidt Fabricius LeConte Erichson Fall Fabricius Solsky Fabricius Fabricius Brown LeConte Horn LeConte Horn LeConte Bedel LeConte Brown LeConte Horn Robinson Gyllenhal Sturm Brown Haldeman Braham Solsky Cartwright Sturm Brown Herbst Herbst Mulsant Linnaeus Motschulsky Reitter Cartwright Beauvois Horn LeConte LeConte Fall Linnaeus Schmidt Melsheimer Herbst Fabricius Herbst Semenov Fabricius Cartwright Say VanDyke Brown Brown Fabricius LeConte 1887 1843 1887 1840 1840 1775 1858 1848 1907 1801 1876 1792 1792 1928 1850 1872 1887 1857 1911 1857 1928 1878 1870 1946 1808 1805 1929 1848 1790 1876 1957 1800 1928 1789 1783 1842 1761 1866 1892 1944 1805 1870 1878 1878 1932 1758 1907 1845 1863 1789 1787 1798 1898 1781 1944 1835 1918 1929 1928 1775 1878 18 7 1 5 5 11 1 1 12 3 2 6 6 2 5 1 14 1 57 4 2 2 24 14 1 1 8 1 3 260 2 7 6 2 10 15 1 1 1 2 2 18 24 1 3 3 3 158 1 148 29 8 5 2 1 23 4 6 2 19 5 8 11 nea nea nea neo neo neo nea neo nea nea neo neo nea nea nea nea nea nea neo nea nea nea nea nea neo neo nea nea nea neo nea neo nea neo neo neo neo neo neo nea nea nea nea nea nea nea nea nea nea nea neo neo neo neo neo nea nea nea nea nea neo nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae 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Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius Ataenius sphacelatus stercorosus sticticus stuessyi sturmi subterraneus talpoidesi tasmaniae tenellus tenuistriatus terminalis testaceiventris thermicola thomomsi tomentosus troglodytes urostigma vandykei varians vittatus walshi wickhami dunnianus brevicruris castaneus sp. tridentatus carolinus cribrosus kolbei fulginea singularis sp. abditus abditus alternatus alternatus apicalis brevis californicus cognatus confertus cylindrus desertus duncani fattigi gracilis haroldi hirsutus imbricatoides imbricatus imbricatus inquisitus miamii morator n.spp. nugator ovatulus picinus picinus picipes platensis puncticollis Sturm Melsheimer Panzer Gordon Harold Linnaeus Brown Hope Say Horn Say Fall Sturm Brown Muller Hubbard Harold Barrett Duftschmid Say Horn Brown Rivers Cartwright Melsheimer 1805 1845 1798 Say VanDyke LeConte Kraatz Burmeister Bates 1823 1928 1835 1895 1847 1888 Haldeman Brown Melsheimer Melshiemer Hinton Fall Horn LeConte Fall Horn Horn Cartwright Cartwight Melsheimer Steinheil Horn Schmidt Melsheimer Melsheimer Horn Cartwright Harold 1848 1932 1844 1844 1937 1930 1887 1858 1909 1871 1871 1974 1948 1844 1872 1871 1909 1844 1844 1887 1934 1869 Harold Horn Harold Harold Fleutiaux and Salle Blanchard LeConte 1880 1871 1867 1867 1889 1843 1868 1897 1758 1928 1846 1823 1887 1823 1932 1800 1928 1776 1894 1862 1931 1805 1825 1870 1928 1889 1944 1846 1 72 6 1 6 59 5 3 1 4 15 1 9 4 3 1 2 8 12 89 10 1 4 2 6 9 10 6 5 1 11 18 1 36 7 183 183 10 2 7 46 6 34 32 1 23 269 5 37 1 8 91 6 7 25 3 3 1 17 33 44 61 neo nea neo nea neo nea nea aus nea nea nea nea neo nea neo nea neo nea neo nea nea nea nea nea nea nea nea nea nea afr neo neo neo nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea neo neo nea nea nea neo nea neo nea nea neo neo nea nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae 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Boreocanthon Boreocanthon Boreocanthon Boreocanthon Boreocanthon Boreocanthon Bothynus Bothynus Bothynus Bothynus Bothynus Bothynus Bothynus Brahmina Bubas Bubas Caccobius Caccobius Caelorrhina Caelorrhina salutator salutator saxatilis scutellaris semipilosus sp. sp. spp. spretulus stephani stercorator strigatus strigicauda terminalis texanus texanus tuberculatus histeroides illaesum lecontei robustum subquadratum texanus tigrina centaurus ciliatus variabilis echinicollis hollowayorum inopinus atra campestris conspurcata lineolata orientalis onitoides anneae concavum deceptum inclinatum keithi nanum ebenus lecontei melanus praticola probus puncticollis simplex dasypleurus entellus laevicollis selanderi sp. spp. spp. dilaticolis bison bubalus mundus schreberi relucens superba Fabricius),? Fabricius),??? Cartwright Harold VanDyke 1944 1867 1928 Haldemann Cartwright Fabricius Say Bates Chevrolat Harold Harold Schmidt Weber Harold Harold Harold Harold Robinson Olivier Fabricius Solier Philippi Howden Howden Howden Burmeister Latreille Harold Fischer Waterhouse Harold Howden Howden Howden Howden Howden Howden Say Harold Robinson LeConte Germar LeConte LeConte Germar Serville Bates Cartwright 1848 1974 1775 1823 1887 1864 1874 1874 1911 1801 1868 1875 1868 1868 1948 Ballion Linnaeus Olivier Menetries Linnaeus Bates Gerstaecker 1870 1767 1811 1838 1767 1881 1883 1775 1851 1861 1842 1804 1878 1823 1875 1868 1823 1870 1948 1859 1824 1866 1857 1824 1825 1888 1959 1 1 5 1 18 1 2 12 35 1 2 207 2 4 1 28 1 36 2 25 11 4 1 2 18 1 1 1 1 4 11 2 1 10 5 1 1 1 1 1 1 17 2 1 13 19 5 73 4 4 1 2 2 7 18 1 6 1 1 16 1 4 neo neo nea neo nea neo neo nea nea neo nea nea neo neo nea neo nea neo nea neo neo nea afr afr neo neo aus aus aus neo pal pal pal ori neo aus aus aus aus aus aus nea nea nea nea nea nea nea neo neo neo neo neo neo neo pal pal pal pal pal afr afr Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae 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Canthon Canthon Canthon Canthon Canthon Canthon Canthon Cartwrightia Catharsius Catharsius Catharsius Celidota Cetonia Cetonia Cetonia Cetonia concinna splendens inconstans sobrina exiguus exiguus aeneolum ardens aurifex breve haroldi leucopterum lucidum planovultum sp. tenebrosum affinis angularis angustatus bispinatus callosus chalcites chevrolati coeruleicollis conformis curvipes curvodilatum cyanellus cyanellus depressipennis dives fastuosus forreri fulgidus humectus humectus imitator lamprimus lividus moniliatus mutabilis nigricornis pilularius quinquemaculatus rugosus septemmaculatus smaragdulus speculifer spp. triangularis tristis vigilans violaceus viridis intertribalis molossus pithecius sp. stephens aurata aurata aurata aurata Blanchard Burmeister Brookes Sharp Boheman Bohemann Harold Bates Bates Germar P.deBorre Howden and Young Harold Howden and Young 1850 1844 Howden and Young Fabricius Harold Harold Robinson Harold Halderman Harold Blanchard Harold Harold Schaum LeConte LeConte LeConte Harold Harold Bates Redtenbacher Say Say Brown Bates Blanchard Bates Lucas Say LeConte Castelnau Blanchard Latreille Fabricius Castelnau 1981 1801 1868 1867 1941 1868 1843 1868 1843 1868 1868 Drury Harold LeConte Olivier Beauvois Islas Linnaeus Fabricius 1770 1862 1858 1789 1805 1958 1758 1775 Gory and Percheron Linnaeus Reitter Fieber Reitter 1833 1761 1896 1831 1896 1876 1860 1860 1867 1887 1887 1824 1886 1981 1867 1981 1859 1859 1859 1868 1868 1887 1867 1832 1832 1887 1843 1887 1857 1823 1758 1840 1868 1807 1781 1840 2 1 1 4 6 6 1 1 1 6 4 1 1 2 2 2 1 2 2 1 3 25 1 2 6 4 3 3 10 4 2 4 1 4 26 44 42 1 2 1 1 31 124 1 7 13 5 2 8 19 7 21 4 4 11 9 8 1 2 52 1 3 9 neo neo aus aus afr afr neo neo neo neo neo neo neo neo neo neo neo neo neo nea neo nea neo neo neo neo neo neo nea nea neo neo neo neo neo nea nea neo neo neo neo nea nea neo neo neo neo neo nea neo neo nea neo nea neo ori ori ori afr pal pal pal pal Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Cetonia Cetonia Cetonia Cetonia Cetonia Cetonia Cetonia Cetonia Cetonia Chalcocopris Chalcocrates Chalcoplethis Chalcosoma Chalcosoma Chalcosoma Chalepides Chalepides Chasmatopterus Chasmatopterus Cheiroplatys Chelorrhina Chelorrhina Chioneosoma Chioneosoma Chioneosoma Chioneosoma Chiron Chiron Chiron Chironitis Chironitis Chironitis Chlorocala Chlorocala Chlorocala Chlorocala Chlorochiton Chnaunanthus Chnaunanthus Chrysina Chrysina Chrysophora Clinteria Clinteroides Cnemida Cnemida Coelosis Coelosis Coelosis Coelosis Coelosis Coenoyncha Coenoyncha Copris Copris Copris Copris Copris Copris Copris Copris Copris Copris aurata aurata aurata aurata chrysosoma magnifica pilifera roelofsi viridiopaca hesperus sp. velutipes atlas atlas sp. barbatus spp. hirtulus vilosulus borealis polyphemus savagei gorilla mucosum porosum reitteri cylindrus grandis puncticollis furcifer hungaricus irroratus africana africana sp. viridicyanea suturalis chapini flavipennis amoena macropus chrysochlora spilota permutans lacerata retusa bicornis biloba sp.1 sp.2 sp.3 fusca testacea arizonensis boucardi capensis costaricensis eudamas fidius fricator hispanus howdeni incertus Reitter Reitter Reitter Reitter Reitter Ballion Motschulsky Harold Motschulsky Olivier 1896 1896 1898 1896 1896 1870 1860 1880 1860 1789 Arrow Linnaeus Erichson 1900 1758 1834 Fabricius 1787 Illiger Illiger Carne Fabricius Harris Brenske Semen Fischer Semen Fabricius Gory Harold Rossi Herbst Rossi Drury Harold 1803 1803 1957 1781 1844 1886 Pallas Fabricius Saylor Horn Sturm Francillon Latreille Gory and Percheron Burmeister Germar Fabricius Leske Linnaeus 1805 1775 1937 1867 1843 1795 1811 1833 1847 1824 1801 1779 1767 McClay Cazier Schaeffer Harold Waterhouse Gahan Olivier??? Olivier Fabricius Linnaeus Matthews and Halffter, Say 1943 1937 1906 1869 1891 1894 1824 1887 1798 1829 1862 1792 1789 1792 1773 1878 1789 1787 1764 1835 3 2 4 1 1 4 1 5 1 1 1 1 5 2 2 16 3 1 1 1 4 3 1 1 1 1 1 14 2 5 3 1 1 1 1 1 3 5 1 1 11 8 1 2 1 10 28 10 1 2 1 2 1 13 1 2 1 1 1 44 4 1 1 pal pal pal pal pal pal pal pal pal neo aus neo ori ori ori neo neo pal pal aus afr afr pal pal pal pal pal afr afr pal pal pal afr afr afr afr aus nea nea neo neo neo pal afr neo neo neo neo neo neo neo nea nea nea neo afr neo neo ori nea pal nea neo Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Copris Copris Copris Copris Copris Copris Copris Copris Copris Copris Copris Copris Copris Copris Copris Copris Copris Copris Copris Coproecus Coptomia Coscinocephalus Coscinocephalus Cotalpa Cotalpa Cotalpa Cotalpa Cotalpa Cotalpa Cotinis Cotinis Cotinis Cotinis Cotinis Cotinis Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Cremastocheilus Ctenotis incertus inemarginatus jacchus klugi klugi laeviceps lecontei lugubris lugubris lunaris macer minutus moechus nevinsoni ochus remotus repertus sacontala tullius hemisphaericus prasina cribrifrons cribrifrons consobrina consobrina flavida lanigera lanigera subcrinita lebas mutabilis mutabilis nitida pauperula sp. academicus angularis armatus armatus canaliculatus castanea constricticollis crinitis crinitus harrisi hirsutus hirsutus incisus knochi lengi maritimus mentalis mexicanus mexicanus nitens planatus planipes schaumi squamulosus tomentosus tridens variolosus obesa Say Blatchley Fabricius Harold Harold Harold Matthews Boheman Bohemann Linnaeus Peringuey Drury LeConte Waterhouse Motschulsky LeConte Walker Redtenbacher Olivier Guerin Burmeister Schaeffer Schaeffer Horn Horn Horn Linnaeus Casey Wickham Gory and Percheron Gory and Percheron Gory and Percheron Linnaeus Burmeister 1835 1918 1775 1869 1869 1869 1858 1858 1758 1900 1773 1854 1891 1860 1866 1858 1848 1789 1830 1842 1915 1915 1871 1871 1878 1764 1915 1905 1833 1833 1833 1764 1847 2 1 1 4 12 1 5 6 6 19 1 76 1 3 2 3 4 1 47 1 1 17 53 13 30 2 65 7 5 1 8 244 90 1 Krikken LeConte Walker Casey Kirby Knoch Cazier LeConte LeConte Kirby VanDyke VanDyke Casey LeConte Cazier Casey Cazier Schaum Schaum LeConte LeConte Horn LeConte LeConte Warner Casey Kirby Burmeister 1982 1857 1866 1915 1827 1801 1940 1874 1874 1827 1918 1918 1915 1853 1940 1915 1940 1841 1841 1853 1863 1885 1853 1858 1985 1915 1826 1855 3 2 3 6 12 51 1 1 3 15 1 2 1 5 1 2 1 2 5 3 1 3 3 1 2 2 3 2 neo nea afr neo neo neo neo neo neo pal afr nea neo ori ori nea ori ori nea aus afr nea pal neo nea nea nea nea nea neo neo nea nea neo neo neo nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea neo Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Cyclidinus Cyclidius Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyclocephala Cyphochilus Cyprolais Cyriopertha Cyriopertha Dasyvalgus Dasyvalgus Deltochilum Deltochilum Deltochilum Deltochilum Deltochilum Deltochilum Deltochilum Deltochilum Deltochilum Democrates Dendropemon Dialytellus Dialytellus Dialytes Dialytes Dialytes Dialytes Dialytes Diaphonia Dichelonyx Dichelonyx lugubris sp. amazonica borealis brittoni carbonaria castanea castaniella cubana freudei helavai herteli hirta longula lunulata lurida mafaffa melanocephala melanocephala modesta nigricollis olivieri ovulum parallela pasadenae prolongata puberela putrida seditiosa signaticollis sinaloe spp. stictica testacea tetrica villosa farinosus hornimani glabra glabra formosanis spp. amazomicum brasiliense denitpes furcatum gibbosum icarus rubripenne scabriusculum tesselatum burmeisteri convexus dialytoides humeralis criddlei n.sp. striatulus trunctatus ulkei mniszechi albicollis backii Westwood 1874 Fabricius Arrow Endrodi Arrow Olivier Bates Chapin Endrodi Endrodi Endrodi LeConte LeConte Burmeister Bland Burmeister Fabricius Fabricius Burmeister Burmeister Arrow Bates Casey Casey Arrow LeConte Burmeister LeConte Burmeister Howden and Endrodi 1781 1911 1964 1911 1789 1888 1932 1963 1975 1964 1861 1863 1847 1863 1847 1775 1775 1847 1847 1911 1888 1915 1915 1902 1873 1847 1863 1847 1966 Burmeister Burmeister Burmeister Blanchard Waterhouse Bates Semenov Reitter Moser 1847 1847 1847 1846 1867 1877 1890 1890 1915 Bates Castelnau Eschscholtz Castelnau Fabricius Olivier Gory Bates Bates Reiche Harold Fall LeConte Brown,??? 1887 1840 1822 1840 1775 1789 1831 1887 1870 1852 1869 1907 1878 Say Melsheimer Horn Janson Burmeister Barrett 1825 1844 1875 1873 1855 1933 2 1 1 47 1 1 3 1 2 2 1 2 17 36 26 107 10 5 156 2 4 5 89 31 77 1 3 2 12 6 1 144 1 5 6 2 1 2 1 1 2 14 2 1 1 1 19 2 1 1 1 1 1 11 2 2 3 19 23 4 2 52 15 afr neo neo nea neo neo neo neo neo neo neo neo nea nea neo nea neo neo nea neo nea neo pal nea nea neo nea neo nea neo neo neo neo neo nea pal afr pal pal ori ori neo neo neo neo nea neo neo neo neo neo nea nea nea neo nea nea nea aus nea nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dichelonyx Dicheros Dichotomius Dichotomius Dichotomius Dichotomius Dichotomius Dichotomius Dichotomius Dichotomius Dichotomius Dichotomius Dichotomius Dichotomius Dichotomius Dichotomius Dichotomius Dicranocephalus Dicronorhina Dicronorhina Didactylia Didactylia Didactylia Diloboderus Dilochrosis Dinacoma Dipelicus Diphucephala Diplognatha Diplognatha Diplognatha Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis canadensis crotchi decolorata diluta elongata elongatula fulgida fuscula lateralis linearis muscula nana pusilla sp. subvittata sulcata sulcata testacea truncata vaga valida vicina vicina burmeisteri anaglypticus ascanius carolinus carolinus carolinus carolinus colonicus fimbriatus glaucus longiceps mormon nisus semiaeneus sexdentatus torulosus bowringi derbyana oberthueri knausi sp. sp. abderus balteata caseyi optatus colaspidoides gagates gagates subaenea abnormis aequalis ambigua angularis angustula arizonica atlantis atramentaria barbarae belfragei Horn Horn Fall Fall Fabricius Schonherr LeConte LeConte Fall Gyllenhal Fall Fall LeConte LeConte LeConte Saylor Kirby LeConte Fall LeConte Fall Hopping Schoch Mannerheim Harold Linnaeus Linnaeus Say Say Say Harold Harold Taschb. Ljungh. Olivier Germar Luderwaldt Eschscholtz Pascoe Westwood Deyrolle Fall Sturm Vollenhoven Blaisdell Sharp Gyllenhal Forster MacLeay Duviv Fall Cazier Fall LeConte Moser Schaeffer Fall Bates Vaurie and Cazier Fall 1876 6 nea 1876 5 nea 1901 11 nea 1901 16 nea 1792 165 nea 1817 12 nea 1856 45 nea 1856 6 nea 1901 1 nea 1817 3 nea 1901 9 nea 1901 1 nea 1856 11 nea 5 nea 1856 47 nea 1856 2 nea 1933 9 nea 1837 5 nea 1856 26 nea 1901 5 nea 1856 3 nea 1901 10 nea 1931 1 nea 1898 2 ori 1829 5 neo 1869 4 neo 1 neo 40 nea 21 nea 1835 2 neo 1835 1 nea 1869 5 neo 1869 12 neo 1870 4 neo 1799 5 neo 1789 10 neo 1824 4 neo 1 neo 1822 2 neo 3 ori 1843 2 afr 1876 3 afr 1905 5 nea 1 neo 3 neo 1826 12 neo 1871 2 aus 1930 5 nea 1875 1 aus 1817 1 aus 1771 14 afr 1838 4 afr 1891 1 afr 1909 2 nea 1940 2 nea 1909 2 nea 1856 59 nea 1918 4 neo 1907 4 nea 1909 18 nea 1888 5 neo 1955 1 neo 1909 115 nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis bidentata boops brevicollis brevicollis brevicornis brevidens carbonata chiricahuae chiricahuae conformis conformis costulata cribratella cribulosa cribulosa dentella dubia fimbriata frondicola fulva harperi haydeni hebes hirsuta illustris ingenua jamaicensis knausi knausii languida latispina liberta magna magna marginicollis marginicollis maura mentalis mimosae mimosae moerens muricata mus obscura pacata parallela parvicollis pinalica planidens poropyge punctata punctaturugosa rosae rudis rufa rufiola sierrae simplex sordida sp. pubereagroup sp. trapeziferagroup spp. statura LeConte Bates LeConte LeConte Cazier LeConte LeConte Fall Fall Fall Fall Fall Bates LeConte LeConte Fall LeConte Fall Say LeConte Blanchard LeConte Bates Vaurie Fall Fall Cazier Schaeffer Schaeffer LeConte Vaurie Germar Bates Bates Fall Fall Fall Fall Fall Fall LeConte Schaeffer Fall LeConte LeConte Fall Fall Fall Fall Bates LeConte Blanchard Vaurie and Cazier LeConte Linell Fall Fall Blanchard Say Cazier 1856 4 nea 1888 5 nea 1856 6 nea 1856 57 nea 1940 1 nea 1856 1 nea 1856 4 nea 1909 11 nea 1909 32 nea 1909 1 nea 1909 5 nea 1909 1 nea 1889 3 neo 1856 239 nea 1856 227 nea 1909 6 nea 1856 2 nea 1909 5 nea 1825 28 nea 1856 1 nea 1850 20 nea 1856 7 nea 1888 1 neo 1958 4 neo 1909 1 nea 1909 25 nea 1952 6 neo 1907 12 nea 1907 2 nea 1878 12 nea 1960 6 neo 1824 11 nea 1888 4 nea 1888 7 nea 1909 14 nea 1909 144 nea 1909 6 nea 1909 14 nea 101 nea 1909 4 nea 1856 9 nea 1907 1 nea 1932 1 neo 1856 24 nea 1856 34 nea 1909 3 nea 1909 1 nea 1932 7 nea 1909 1 nea 1887 2 neo 1856 3 nea 1850 22 nea 1955 1 neo 1859 1 nea 1896 1 nea 1909 8 nea 1909 1 nea 1850 4 neo 1825 111 nea 1 neo 5 neo 16 nea 2 nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Diplotaxis Dischista Dymusia Dynastes Dynastes Dynastes Dynastes Dynastes Dynastes Dynastes Dyscinetus Dyscinetus Dyscinetus Dyscinetus Dyscinetus Dyscinetus Dyscinetus Dyscinetus Dyscinetus Endroedyantus Enema Enema Enema Eophileurus Epadoretus Epectinaspis Epectinaspis Epirinus Epirinus Eremazus Erioscelis Eucanthus Euchirus Euchirus Euchirus Euchirus Euchirus Euchloropus Eucranium Eudicella Eudicella Euheptaulacus Euheptaulacus Euheptaulacus Euparia Euparia Eupatorus Euphoria Euphoria Euphoria Euphoria Euphoria subangulata subangulata subangulata subcostata sulcatulus tarsalis texana thoracica trapezifera trementina tristis truncatula cincta cyanea granti hercules hercules hercules hyllus neptunus tityus dubius gagates morator picipes picipes rugifrons sp. spp. spp. paradoxus pan pan sp. chinensis reitteri mexicanus opacicollis silvestris validus unistriatus emarginata impressus dupontianus longimanus macleayi parryi sp. laetus arachnoides gralli morgani carinatus sus villosus castanea spp. hardwickii basalis biguttata candezei canescens dimidiata Blanchard LeConte Schaeffer Blanchard Fall Schaeffer LeConte Fall Bates Fall Kirby LeConte DeGeer Olivier Horn Linnaeus Verril Scopoli Chevrolat Quensel Linnaeus Olivier Burmeister Fabricius Burmeister Burmeister Burmeister Scholtz and Howden Fabricius Fabricius 1850 1856 1907 1850 1909 1907 1856 1909 1887 1909 1837 1856 1778 1789 1758 1906 1769 1843 1817 1763 1789 1847 1798 1847 1847 1847 1775 1787 Falderman Semenov Burmeister Bates Cambefort Peringuey Mulsant Mannerheim Howden Burmeister Linnaeus Hope Gray 1835 1890 1844 1888 Fabricius Brulle Buquet White Germar Herbst Gyllenhal Lepeletier and Serville 1801 1834 1836 1839 1824 1783 1808 1828 Hope Gory and Percheron Gory and Percheron Janson Gory and Percheron Gory and Percheron 1831 1833 1833 1875 1833 1833 1901 1854 1829 1 93 6 44 6 2 11 12 71 2 93 63 1 1 46 9 1 1 2 2 68 149 2 150 17 24 6 3 72 176 1 11 6 10 1 1 8 1 2 1 1 33 1 1 1 4 2 2 7 1 3 2 2 4 1 14 3 4 36 3 1 2 2 nea nea neo nea nea nea nea nea neo nea nea nea afr afr nea neo neo neo neo neo nea neo neo nea neo nea neo neo nea afr neo neo neo pal pal neo neo afr afr pal neo nea ori ori ori ori ori ori neo afr afr pal pal pal nea neo ori neo neo neo neo neo Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Euphoria Euphoria Euphoria Euphoria Euphoria Euphoria Euphoria Euphoria Euphoria Euphoria Euphoria Euphoria Euphoria Euphoria Euphoria Euphoria Euphoria Euphoriopsis Eupoecila Europtron Eutheola Eutheola Eutheola Eutheola Eutheola Eutheola Exolontha Gametis Gametis Gametis Gametis Gametis Gametis Gametis Gametis Gametis Gametis Geniates Geniates Genuchinus Geopsammodius Geotrogus Glycyphana Glycyphana Glycyphana Glycyphana Glycyphana Glycyphana Glycyphana Glycyphana Glycyphana Glycyphana Glycyphana Gnathocera Gnathocera Gnorimella Gnorimus Gnorimus Gnorimus Gnorimus Goliathus Goliathus Goliathus fulgida fulgida fulgida fulgida herbacea histrionica inda inda kernii lineoligera pulchella sepulchralis sepulchralis sepulchralis sepulchralis sp. subtomentosa hirtipes australasiae confusum bidentata humilis sp. sp. spp. subglabra mandarinen jucunda jucunda jucunda jucunda jucunda versicolor versicolor versicolor versicolor versicolor bornelli spinolae ineptus hydropicus gabalus cuculus fulvistemma magnifica roquesi rubripes rubroscutellaris sp. sp. nr. mollendorfi spp. stolata tibialis trivittata trivittata maculosa nobilis octopunctatus octopunctatus subopacus cacicus goliatus orientalis Fabricius Casey Fall Fall Olivier Thomas Linnaeus Klages Haldeman Blanchard Gory and Percheron Fabricius Casey Gory and Percheron Gory and Percheron Mannerheim Horn Donovan Marseul? Burmeister LeConte Schaeffer Fairmaire Falderman Gory and Percheron Hope Reitter Reitter Arrow Arrow Arrow Arrow Arrow Camerano Burmeister Horn Horn Fairmaire Burmeister Motschulsky Moser Bourgin Moser Mohnike Fabricius Moser Swederus Swartz Knoch Linnaeus Linnaeus Linnaeus Motschulsky Voet Drury Moser 1775 49 nea 1915 13 nea 1905 44 nea 1905 8 nea 1789 10 nea 1878 32 nea 1760 101 nea 1894 1852 95 nea 1850 2 neo 1833 17 neo 1801 227 nea 1915 17 nea 1833 4 neo 1833 12 nea 2 nea 1837 3 neo 1880 2 nea 1805 1 aus 1878 1 pal 1847 66 neo 1856 23 nea 2 pal 18 nea 64 neo 1909 6 nea 1 ori 1835 16 ori 1833 5 ori 1831 3 ori 1896 2 ori 1898 1 ori 1910 2 ori 1910 1 ori 1910 1 ori 1910 2 ori 1910 1 ori 1894 2 neo 1844 9 neo 1885 2 nea 1887 14 nea 1860 2 pal 1842 8 ori 1860 5 pal 1906 4 ori 1916 2 ori 1906 2 ori 1873 1 ori 1 afr 1 ori 19 ori 2 aus 1913 24 ori 1789 2 afr 1817 3 afr 16 nea 1758 9 pal 1758 1 pal 1758 3 pal 1860 1 pal 1779 4 afr 1770 10 afr 1909 1 afr Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Goliathus Goliathus Goliathus Golofa Golofa Golofa Golofa Golofa Gromphas Gromphas Gronocharus Gronocharus Guatemalica Gymnetis Gymnetis Gymnetis Gymnetis Gymnetis Gymnetis Gymnetis Gymnetis Gymnetis Gymnetis Gymnetis Gymnetis Gymnetis Gymnetis Gymnopleurus Gymnopleurus Gymnopleurus Gymnopleurus Gymnopleurus Gymnopleurus Gymnopleurus Gymnopleurus Gymnopleurus Gymnopleurus Gymnopleurus Gymnopleurus Gymnopleurus Gymnopleurus Gymnopyge Gymnopyge Hadrosticta Haplidia Haplidia Heliocopris Heliocopris Heliocopris Heliocopris Heliocopris Heliocopris Heliocopris Hemictenius Hemictenius Hemictenius Heptophylla Heterogomphus Heterogomphus Heterogomphus Heterogomphus Heteronychus Heteronychus orientalis regius sp. eacus pelegon pizarro porteri spp. aeruginosa lacordairei autumnalis multispinosus marginicollis chalcipes cinerea cretacea holosericea insculpta irregularis maculosa pantherina rufolatris sallei sp. sp. nr sallei tigrina undulata aciculatus coriarius coriarius coriarius cyaneus geoffroyi miliaris mopsus mopsus sinuatus smaragdifer sturmi sturmi virens hirsuta hopliaformis viridiflua graeca transversa bucephalus dominus gigas hamadryas japetus neptunis neptunis nigrociliatus opacipes simplicitarsis picea chevrolati dilaticollis pauson schonherri arator sp. Moser Klug 1909 1835 Burmeister Burmeister Hope Hope 1847 1847 1837 1837 Perty Brulle Schaeffer Howden Burmeister Gory and Percheron Gory and Percheron LeConte Voet Kirby Gory and Percheron Olivier Burmeister Illiger Schaum 1830 1834 Gory and Percheron Vigors Gebler Mulsant Mulsant Fischer Fabricius Fuessely Fabricius Pallas Pallas Olivier Walker MacLeay Duft Erichson Cazier Linell Kraatz Kraatz Fabricius Fabricius Bates Olivier Fabricius Klug Boheman Bohemann Reitter Reitter Reitter Motschultsky Burmeister Burmeister Perty Burmeister Fabricius 1833 1825 1841 1842 1842 1821 1798 1775 1775 1781 1781 1789 1858 1821 1805 1843 1939 1896 1892 1882 1801 1775 1868 1798 1775 1855 1857 1857 1897 1901 1897 1857 1847 1847 1830 1847 1775 1967 1842 1833 1833 1863 1779 1818 1833 1789 1842 1800 1849 2 4 3 1 2 3 2 18 1 6 8 4 3 1 6 2 afr afr afr neo neo neo neo neo neo neo nea nea neo neo neo nea 1 1 1 1 1 5 1 1 5 2 2 1 2 1 1 4 1 2 4 1 2 9 1 1 5 16 1 1 2 6 1 4 1 1 1 1 1 1 1 1 15 2 5 5 4 21 neo neo neo neo neo nea neo neo neo neo pal pal pal pal ori pal ori pal pal ori ori pal pal afr nea nea neo pal pal ori ori afr afr afr afr afr pal pal pal ori neo neo neo neo aus afr Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Heterophana Heterorhina Heterorhina Heterorhina Holotrichia Holotrichia Homalopia Homalopia Homalopia Homonyx Homovalgus Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplia Hoplopyga Hoplopyga Hoplopyga Hoplopyga Hoplopyga Hoplopyga Hylamorpha Hylamorpha Hymenoplia Hymenoplia Hypothyce Idutywa Inca Incala Ischiopsopha Ischiopsopha Ischiopsopha Ischiopsopha Ischiopsopha Ischiopsopha Ischiopsopha Ischiopsopha Ischnoscelis Isonychus Isonychus Lachnota Leiopsammodius Leptohoplia Leptothyrea lineatta dohni imperatrix punctatissima diomphalia morosa marginata ruricola spiraeae sp. seticollis angulata aulica aulica aureola callipyge caucasica coerulea dispar equina farinosa graminicola modesta parvula paupera philanthus practicola rufipes sackenii sp. sp. sulphurea trifasciata trivialis brasiliensis liturata miliaris ocellata sp.1 sp.2 elegans elegans cinerascens rugulosa mixta collaris clathratus sp. esmeralda jamesi lucivorax rugata sp. sp. virens wallacei dohrni arizonensis ocellatus henningi malkini testaceipennis perroudi Burmeister Lansberge Mohnike Westwood Bates Waterhouse Fuessley Fabricius Pallas 1842 1883 1871 1842 1888 1875 1775 1775 1773 Beauvois Reitter Linnaeus Fabricius Pallas LeConte Kolenati Drury LeConte LeConte Linnaeus Fabricius Haldeman Krynicki Krynicki Fussley Duftschmid Motschulsky LeConte 1805 1902 1766 1801 1781 1856 1846 1773 1880 1880 1761 1792 1843 1832 1832 1775 1805 1860 1880 Dufour Say Harold Gory and Percheron Olivier Gory and Percheron Gory and Percheron 1843 1823 1869 1833 1789 1833 1833 Burmeister Phillipi ? Rosenhauer Mulsant Howden Burmeister Olivier 1844 Wallace Waterhouse Kraatz Blanchard 1868 1876 1890 1842 Blanchard Thomas Westwood Howden Burmeister Fischer Cartwright Saylor Schaum 1853 1857 1855 1856 1842 1968 1844 1792 1855 1836 1946 1935 1844 2 3 1 4 1 8 1 4 4 2 24 1 2 1 2 63 1 31 11 6 1 2 21 1 1 14 2 1 5 1 3 8 177 3 26 1 3 1 1 3 6 4 1 2 2 2 4 4 2 1 3 1 2 5 2 1 1 9 9 1 3 2 1 afr ori ori ori pal ori pal pal pal neo nea pal pal pal ori nea pal neo nea nea pal pal nea pal pal pal pal ori nea neo pal pal nea nea neo neo neo neo neo neo neo neo pal pal nea afr neo afr pal pal pal pal pal aus pal aus neo nea neo pal nea nea afr Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Leucoserica Leucothyreus Ligyrus Ligyrus Ligyrus Ligyrus Ligyrus Ligyrus Ligyrus Ligyrus Ligyrus Ligyrus Ligyrus Ligyrus Ligyrus Ligyrus Ligyrus Ligyrus Ligyrus Liocola Liocola Liocola Liocola Liocola Liocola Liogenys Lomanoxoides Lomaptera Lomaptera Lomaptera Lomaptera Lomaptera Lomaptera Lomaptera Macraspis Macraspis Macraspis Macraspis Macraspis Macraspis Macrodactylus Macrodactylus Macrodactylus Macrodactylus Macrodactylus Macrodactylus Macrodactylus Macrodactylus Macronota Macronota Macronota Macronota Maladera Maladera Maladera Maladera Martinezia Mausoleopsis Megaceras Megaceras Megaceras Megaceras Megalorhina arenicola kirbyanus cuniculus ebenus fossor gibbosus gibbosus gibbosus gyas neglectus neglectus relictus sallaei sallaei sp. sp. spp. subtropicus villosus brevitarsis brevitarsis catharica lugubris lugubris speculifera palpalis bitubericolis cinnamomea helleri salvadorii satanus simbagensis soror wahnesi chalcea cincta dichroa festiva lucida tetradactyla angustatus fulvescens mexicanus nigricornis? subspinosus thoracicus uniformis uniformis nigritus regia sp. spp. castanea holosericea orientalis renardi excavaticollis amabilis jason morpheus philoctetes spp. harrisi Solsky Macleay Fabricius De Geer Latreille DeGeer LeConte LeConte Erichson LeConte LeConte Say Bates Bates Blatchley Burmeister Lewis Kraatz Bates Herbst Fabricius Swartz Eschscholtz Schmidt Thomas Moser Gestro Schaum Schaum Kraatz Moser Burmeister MacLeay Waterhouse Burmeister Olivier Linnaeus Beauvois Bates Burmeister Moser? Fabricius Kirsch Horn Horn Waterhouse Fabricius Arrow Scopoli Motschulsky Ballion Blanchard Schaum Fabricius Burmeister Olivier Westwood 1876 1 pal 1819 3 neo 1801 46 neo 1774 2 neo 1833 2 neo 1774 244 nea 1847 9 nea 1856 116 nea 1848 15 neo 1847 4 neo 1847 34 nea 1825 37 nea 1888 6 neo 1888 7 nea 1 nea 2 neo 190 1922 4 nea 1847 10 neo 1879 22 afr 1889 1 ori 1890 1 ori 1786 1 ori 1792 3 pal 1817 4 ori 1822 2 neo 1909 1 neo 1878 1 pal 1908 2 pal 1876 1 aus 1912 2 pal 1 pal 1890 3 afr 1906 1 pal 1844 8 neo 1819 4 neo 1881 6 neo 1844 6 neo 1789 5 neo 1771 4 ori 1805 5 nea 1887 6 neo 1855 7 neo 1 neo 1775 108 nea 1885 3 neo 1876 36 nea 1876 46 nea 1841 1 ori 1801 4 aus 4 ori 19 ori 1913 52 nea 1772 12 pal 1857 1 pal 1870 1 neo 1843 2 nea 1844 1 afr 1775 2 neo 1847 7 neo 1789 1 neo 6 neo 1847 3 afr Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae 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gyas mars thersites argentina bicolor villosa umbonata hippocastani hippocastani melolontha pectoralis vulgivagus chinensis junii splendens validipes xanthopus alutaceum laevis pecuarius castanopterus interuptus quinqueplicatus werneri werneri werneri mimicus stanleyi endroedyi peckorum armaticeps bidens australis halii nitidula smithii striata varicolorata xanthosticta zealandica floralis festivus fulvus fulvus militarius nasicornis pallens pallipes sp. aerarius crenatus falcatus humerosus ion sphinx virens appendiculatus sulcator howdeni lerati Linnaeus Chevrolat Fabricius Herbst Reiche LeConte Gillet Guerin Eschscholtz Gory and Percheron Fabricius Comolli Linnaeus Germar Olliff Kirby Duftschmidt Gyllenhal Heller Boisduval Afzeli Burmeister Reiche Hope Say Horn Cartwright Cartwright Cartwright Gordon and Cartwright Janson Howden and Scholtz Howden and Scholtz Fall Horn Given Broun Broun Broun White Given White White LeConte Steven Goeze Fabricius Castelnau Reiche Olivier Fabricius Horn Harold Reiche Wulfen Pallas Olivier Fabricius Lansberge Mannerheim Fabricius Paulian Paulian 1758 1836 1775 1785 1852 1861 1911 1839 1839 1833 1801 1837 1758 1824 1889 1825 1805 1817 1894 1835 1817 1847 1860 1835 1871 1955 1955 1955 1980 1889 1932 1871 1893 1846 1846 1846 1856 1809 1777 1781 1840 1849 1789 1781 1882 1878 1847 1786 1771 1789 1775 1875 1829 1775 21 2 23 1 1 1 1 8 6 2 7 2 36 1 1 1 2 1 2 1 1 1 1 2 3 24 1 1 5 1 1 1 4 2 9 4 3 3 3 3 4 1 4 5 1 15 4 4 4 1 9 2 2 2 1 1 1 2 1 10 2 2 2 neo neo neo neo neo neo neo neo neo afr pal pal pal pal aus ori pal ori aus ori afr aus aus pal nea nea nea neo neo neo afr afr afr nea nea aus aus aus aus aus aus aus aus nea pal pal pal afr afr pal pal nea afr pal ori pal pal afr pal neo neo aus aus Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera 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Onthophagus Onthophagus Onthophagus montrouzieri alluvius amyntas anthracinus ater atramentarius bonasus brevifrons browni castanea chevrolati cochisus coenobita concinnus concinnus crocatus crocatus depressus finschi fissicornis flagrans fracticornis furcatus gazella gazella gazella gazella germanus gibbosus gibbulus granulatus granulatus grossepunctatus haroldi hecate hecate hoepfneri hopfneri knausi knulli koshantschikoffi landolti lecontei lemur marginalis marginatus marginicollis medorensis mexicanus nebulosus nuchicornis nuchicornis oklahomensis orpheus orpheus ovatus pennsylvanicus polyphemi polyphemi pygargus rangifer ruficapillus sibiricus Paulian Howden and Cartwright Olivier Harold Waterhouse Menetries Linnaeus Horn Howden and Cartwright Serville Harold Brown Herbst Castelnau LaPorte Mulsant,??? Mulsant? Harold Harold Steven Reitter Preyssler Fabricius Fabricius Fabricius Fabricius),? Fabricius),?? Linnaeus Reitter Pallas Boheman Bohemann Muller Ball Panzer Brown Harold Harold Brown Howden and Cartwright Reitter Harold Harold Fabricius Gebler Castelnau Harold Brown Bates Reiche Linnaeus Linnaeus Brown Panzer Fabricius Linnaeus Harold Hubbard Howden and Cartwright Motschuskly Klug Brulle Harold 1789 1873 1875 1832 1775 1881 1825 1869 1927 1783 1840 1840 1871 1877 1809 1893 1790 1781 1787 1767 1891 1781 1858 1858 1906 1871 1794 1929 1869 1869 1927 1963 1891 1880 1871 1781 1817 1840 1880 1929 1887 1864 1758 1758 1927 1794 1801 1767 1871 1894 1963 1845 1855 1832 1877 2 1 4 7 2 1 9 1 8 14 2 4 1 7 1 1 13 2 1 1 26 20 27 3 17 17 3 3 2 3 3 6 2 100 11 1 5 1 2 2 5 5 11 1 14 2 12 3 1 12 69 11 15 6 26 42 1 3 1 2 6 1 aus nea pal nea ori pal ori nea nea nea neo nea pal nea nea pal pal nea ori pal pal pal pal neo afr nea nea pal pal pal aus aus pal pal nea nea nea neo nea nea pal nea neo pal pal neo neo nea neo afr pal nea nea nea nea pal nea nea nea pal afr pal pal Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Onthophagus Onthophagus Onthophagus Onthophagus Onthophagus Onthophagus Onthophagus Onthophagus Onthophagus Onthophagus Onthophagus Onthophagus Onthophagus Onthophagus Onthophagus Onthophagus Oplostomus Orizabus Oruscatus Oryctes Oryctes Oryctes Oryctes Oryctes Oryctes Oryctes Oryctes Oryctes Oryctes Oryctes Oryctoderus Oryctomorphus Oryctomorphus Osmoderma Osmoderma Osmoderma Oxygrylius Oxyligyrus Oxyomus Oxysternon Oxysternon Oxysternon Oxythyrea Oxythyrea Oxythyrea Oxythyrea Oxythyrea Oxythyrea Oxythyrea Oxythyrea Oxythyrea Oxythyrea Oxythyrea Oxythyrea Oxythyrea Pachnoda Pachnoda Pachnoda Pachnoda Pachnoda Pachnoda Pachnoda Pachnoda speculifer spp. striatulus striatulus striatulus subaeneus subtropicus tapirus taurus taurus tuberculifrons vacca velutinus velutinus verticicornis vitulus fuligineus clunalis opalescens boas gigas gnu nasicornis nasicornis rhinoceros rhinoceros rhinoceros sp. sp. spp. latitarsis bimaculatus maculicollis eremicola eremita scabra ruginasus nasutus silvestris conspicillatum festivum palaemon amethystina cinctella funesta funesta haemorrhoridalis hildebrandti marginalis pantherina plebeja rubra spp. testaceoguttata tripolitano consentanea interrupta marginata marginella rufa savigny sinuata sp. Solsky 1876 Beauvois Beauvois Beauvois Beauvois Howden and Cartwright Sharp Schreber Schreber Harold Linnaeus Horn Horn Laich Fabricius Olivier LeConte Bates Fabricius Castelnau Mohnike Linnaeus Illiger Linnaeus Linnaeus Linnaeus 1809 1809 1809 1811 1963 1877 Boisduval Guerin Guerin Knoch Scopoli Beauvois LeConte Burmeister Scopoli Weber Linnaeus Castelnau MacLeay Schaum Poda Linnaeus Fabricius Kraatz Swartz Gory and Percheron Kolbe Gory and Percheron 1835 1830 1838 1801 1763 1805 1856 1847 1763 1801 1767 1840 1838 1841 1761 1766 1775 1880 1817 1833 1895 1833 Blanchard Reitter Schaum. Olivier Drury Fabricius DeGeer Gory and Percheron Gory and Percheron 1850 1891 1844 1789 1773 1775 1778 1833 1833 1759 1873 1767 1875 1781 1776 1789 1856 1870 1775 1840 1874 1758 1803 1758 1758 1758 1 22 1 2 78 8 1 16 32 14 23 15 35 1 8 2 4 280 5 3 8 4 12 6 5 12 12 1 7 12 3 8 4 64 5 86 262 1 4 13 106 11 4 23 61 2 3 3 4 1 1 1 11 4 6 2 2 2 1 15 2 6 1 pal neo nea nea nea nea nea neo nea pal nea pal nea nea pal pal afr nea neo afr afr ori pal pal afr ori pal neo afr aus neo neo nea pal nea nea neo nea neo neo neo afr pal pal pal afr afr afr pal afr afr afr afr pal afr afr afr afr afr afr afr afr Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Pachnoda Pachnoda Pachnoda Pachnoda Pachnoda Pachnodella Pachydema Pachydema Pachylomerus Panglaphyra Panotrogus Parabrysopolis Parabrysopolis Paracotalpa Paracotalpa Paragnorimus Paragymnetis Paragymnetis Parastasia Parataenius Parataenius Parathyce Parathyce Parathyce Pedinorrhina Pelidnota Pelidnota Pelidnota Pelidnota Pelidnota Pelidnota Pelidnota Pelidnota Pelidnota Pelidnota Pelidnota Pelidnota Pelidnota Pelidnota Pelidnota Pelidnota Pelidnota Pentodon Pentodon Pentodon Pericoptus Pericoptus Phaeadoretus Phaenomeris Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus sp. sp. spp. stehelini tridentata impressa bucharica hirticollis femoralis duboulayi myschenkowi arizonae chihuahuae puncticollis ursina sambucas hebraica spp. brevipes simulator simulator fieldi harfordi palpalis mediana chalcothorax fulva laevissima ludovici lugubris lutea lutea lutea nitescens notata prasima prolixa pulchella punctata spp. spp. virescens bidens bidens idiota punctatus truncatus comptus magnifica amethystinus amithaon batesi bellicosus bispinus bonariensis cerberus corythus damon daphnis dardanus difformis ensifera faunus Schaum Olivier Goldfuss Reitter Fabricius Kirby Thomas Ballion Ohaus? Bates LeConte Horn Howden Drapiez 1841 1789 1805 1898 1787 1828 1879 1871 1915 1888 1863 1867 1970 1820 LeConte Harold Harold Fall Casey Horn Westwood Perty Blanchard Burmeister Ohaus LeConte Olivier Casey Casey Vigors Blanchard Burmeister Sharp Kirby Linnaeus 1856 1868 1868 1908 1889 1880 1841 1834 1850 1855 1905 1874 1789 1915 1915 1825 1850 1844 1877 1818 1758 Burmeister Pallas Villers Herbst White Fabricius Menetries Hope Harold Harold Harold Olivier Bates Gory Harold Harold Castelnau Harold MacLeay LeConte Germar Fabricius 1844 1771 1789 1789 1846 1787 1849 1835 1871 1875 1868 1789 1887 1844 1869 1863 1840 1863 1819 1847 1812 1775 3 8 9 1 1 2 1 5 2 4 1 5 3 5 17 1 1 3 9 5 116 1 3 9 1 3 2 1 11 16 7 2 18 2 3 1 1 1 176 22 117 32 3 7 6 3 2 1 2 3 11 8 2 1 2 1 1 7 23 1 19 9 3 afr afr afr afr afr afr pal pal afr afr pal nea nea nea nea neo neo neo nea nea nea nea nea nea afr neo neo neo neo nea nea nea nea neo neo neo neo neo nea nea neo neo pal pal pal aus aus afr pal neo nea neo neo neo neo neo neo neo neo neo nea neo neo Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Phanaeus Pharaonus Pharaonus Phileurus Phileurus Phileurus Phileurus Phileurus Phileurus Phileurus Phileurus Phileurus Phileurus Phileurus Phileurus Phileurus Phileurus Phileurus Phileurus Philoscaptus Phobetus Phobetus Phobetus Phobetus Phobetus Phyllognathus Phyllopertha Phyllopertha Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga furcosus hermes howdeni howdeni igneus imperator jasius jasius kirbyi laeviceps lancifer leander melampus meleagris mexicanus milon palaeno prasinus pyrois quadridens sallei sapphirinus septentrionalis splendidulus spp. sulcatus torrens triangularis velutinus vindex lederi semenowi didymus illatus quadrituberculatus sp. sp. 4 sp.1 sp.2 sp.3 sp.5 spp. truncatus truncatus valgus valgus valgus valgus bonariensis centralis comatus mojavus palpalis spp. excavatus horticola massageta acinosa aemula aequalis analis anxia arcta Bates Harold Arnaud Arnaud MacLeay Chevrolet Olivier MacLeay Vigors Harold Linnaeus Waterhouse Harold Blanchard Harold Blanchard Blanchard Harold Bates Say Harold Sturm Pess??? Fabricius 1887 1868 1983 1984 1819 1844 1789 1819 1825 1869 1767 1891 1863 1843 1863 1843 1843 1868 1887 1835 1863 1826 Drury Olsontieff Edmonds Murray MacLeay Reitter Reitter Linnaeus LeConte Beauvois 1770 1924 1994 1856 1819 1888 1887 1758 1854 1806 Beauvois Beauvois Olivier Bates Casey Olivier Burmeister Casey Barrett Barrett Saylor 1807 1807 1789 1888 Forster Linnaeus Kirsch Arrow Horn LeConte Burmeister LeConte Horn 1771 1758 1881 1920 1887 1857 1855 1850 1887 1781 1789 1847 1909 1856 1933 1936 7 2 1 1 41 6 27 3 1 11 84 1 3 6 2 1 6 1 21 1 8 1 43 21 4 1 1 2 118 1 1 2 26 1 4 1 3 1 1 1 10 1 25 10 1 6 16 7 1 1 11 1 2 7 26 2 2 7 4 1 118 2 neo neo neo neo nea neo neo neo neo neo neo neo neo neo neo neo neo neo neo nea neo neo neo neo neo neo nea nea neo nea pal pal neo nea neo nea neo neo neo neo neo neo neo nea neo neo nea nea neo nea nea nea nea nea pal pal pal neo nea nea neo nea nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae 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Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga arizona arkansana balia barda batillifer beckeri bellamyi bilobatata bilobatata bipartita brevidens bruneri calceata cephalicus chapini chiriquina citri clemens clypeata congrua corrosa crassissima crenulata cribrosa cribrosa crinata crinita cushmani debilis delata densepunctata diffinis dispar disparilis disparilis divertens drakei duncani duncani elizoria ephilida ephilida epigaea errans falsa farcta fervida floridana forbesi forsteri foxii fragilipennis fraterna fucata fulviventris fusca futilis gaigei georgiana glabberima glabricula gracilis gracilis Saylor Schaeffer Say Horn Bates Moser Warner and Moron Saylor Saylor Horn Bates Chapin LeConte Horn Saylor Bates Smyth Horn Horn LeConte LeConte Blanchard Froelich LeConte LeConte Smith Burmeister Say LeConte Horn Moser Blanchard Burmeister Horn Horn Bates Kirby Barrett Barrett Saylor Say Reinhardt Wickham LeConte LeConte LeConte Fabricius Robinson Glasgow Burmeister Davis Blanchard Harris Horn Moser Froelich LeConte Sanderson Horn Blanchard LeConte Burmeister Glasg. 1940 1906 1825 1887 1992 1939 1939 1887 1856 1940 1917 1887 1887 1856 1856 1850 1792 1853 1853 1889 1855 1940 1856 1887 1850 1855 1878 1878 1933 1933 1937 1825 1903 1860 1856 1856 1781 1838 1916 1850 1920 1850 1842 1887 1792 1850 1948 1850 1856 1855 1925 1 9 43 10 1 20 1 55 165 30 1 27 10 4 9 9 7 4 1 8 4 36 71 8 49 6 19 1 31 26 6 4 8 3 7 2 41 1 42 33 37 2 3 4 12 12 18 3 9 39 2 35 5 2 86 58 nea nea nea nea neo nea nea nea nea nea neo nea nea nea nea neo neo nea nea nea nea nea nea nea nea nea nea nea nea nea neo nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea neo nea nea neo nea nea 1 73 4 28 12 nea nea nea nea nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae 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Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga granti granti hirsuta hirticula hirtiventris hoogsraali hornii huachuca huachuca idonea ignava ilicis ilicis impar implicata incuria inepta infidelis inflexa inversa jonesi juvenilis karlsioei kentuckiana knochii koehleriana lanceolata lanceolata latidens latidens latifrons lenis lenis lobata lodingi longispina longitarsa lota luctuosa luginbilli marginalis micans micans misteca mucorea neomexicana? nitida obsoleta opacicollis opacita orosina pallida parvidens parvisetis paternoi plena plenus pleroma postrema praetermissa profunda prolixa prunina Saylor Saylor Knoch Knoch Horn Saylor Smith Saylor Saylor Sanderson Horn Knoch Knoch Davis Horn Sanderson Horn Horn Barrett Horn Sanderson Fall Linell Richter Gyllenhal Saylor Say Say Schaeffer Schaeffer LeConte Horn Horn Fall Sanderson Smith Say Luginbill Horn Saylor LeConte Knoch Langston Bates LeConte Saylor LeConte Blanchard Horn Reinhard Moser Horn LeConte Bates Glasgow Fall Fall Reinhard Horn Horn Blanchard Bates LeConte 1940 1940 1801 1801 1887 1889 1940 1940 1948 1887 1801 1801 1920 1887 1942 1887 1887 1935 1887 1932 1898 1937 1817 1940 1825 1825 1906 1906 1856 1887 1887 1908 1939 1889 1824 1928 1887 1941 1856 1801 1924 1888 1856 1940 1856 1850 1878 1939 1885 1856 1888 1925 1932 1932 1940 1887 1887 1850 1888 1856 2 8 17 89 9 2 8 24 32 8 28 2 45 2 17 1 2 16 1 15 2 1 2 2 7 4 6 27 6 25 64 1 19 6 2 23 24 2 18 1 37 32 3 2 44 8 10 2 2 2 1 1 8 1 1 5 3 4 1 8 9 2 17 nea nea nea nea nea neo nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea nea neo nea nea nea neo nea nea neo nea nea neo nea nea nea nea nea nea nea neo nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae 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Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga Phyllophaga prununculina pusillidens pygidialis quercus ravida rolbakeri rorulenta rossi rubiginosa rubricosa rugosa schaefferi scoparia sequoiana servicata setifera sonora sp. sp. graciliscomplex sp. nrtimidus sp. setidorsisgroup sp.sonora/pallida spp. testaceipennisgroup straminea submucida subpruinosa subtonsa suttonana sylvatica tarsalis tarsalis tarsalis taxodii tecta temora testaceipennis texensis timida timida timida timidus torta transversicollis trichodes tristis tristis tristis trochanter trochanter trochanter tusa ulkei uniformis vanalleni vandykei vehemens vetula vetula vexata villifrons wickhami youngi zeteki Burmeister Fall Schaeffer Knoch Blanchard Saylor Burmeister Saylor LeConte Reinhard Melsheimer Saylor LeConte Saylor Blanchard Burmeister Saylor Bates LeConte Casey LeConte Reinhardt Sanderson Schaeffer Schaeffer Schaeffer Langston Cartwright Saylor Moser Saylor Horn Horn Horn Horn LeConte Moser Bates Fabricius Fabricius Reinhard Saylor Saylor Saylor Horn Smith Blanchard Schaeffer Saylor Horn Horn Horn Horn LeConte Saylor Cartwright Saylor 1855 1937 1906 1801 1850 1855 1939 1856 1939 1846 1937 1856 1936 1885 1939 1856 1884 1856 1942 1908 1908 1908 1924 1944 1921 1940 1878 1878 1878 1856 1890 1781 1781 1939 1940 1940 1940 1887 1888 1850 1927 1887 1887 1887 1885 1856 1940 1935 40 10 19 26 2 2 11 2 11 4 38 7 21 4 5 1 11 1 2 3 8 13 4 2 37 5 4 1 5 2 12 268 6 2 2 1 1 2 4 8 8 44 1 2 1 77 2 1 11 24 1 10 13 10 1 6 24 130 2 8 8 4 2 nea nea nea nea nea neo neo nea nea nea nea nea nea nea neo nea nea nea neo neo nea neo neo nea nea nea nea nea nea nea nea nea nea neo neo nea nea nea nea nea nea neo neo nea nea nea nea nea nea nea nea nea nea neo nea nea nea neo nea nea nea neo Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Pimelopus Plaesiorrhina Plaesiorrhina Plaesiorrhina Platychelus Platygenia Platytomus Platytomus Platytomus Platytomus Platytomus Plectris Plectrodes Plectrone Pleurophorus Pleurophorus Pleurophorus Pleurophorus Plusiotis Plusiotis Plusiotis Plusiotis Plusiotis Podischnus Podolasia Poecilopharis Poecilopharis Poecilophila Poecilophila Poecilophilides Polybaphes Polybaphes Polybaphes Polybaphes Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polyphylla Polystigma Popillia Popillia Popillia Popillia Popillia Popillia Popillia Popillia Popillia dubius recurva recurva watkinsiana dimidiatus barbata atlanticus caelicollis longulus micros notialis aliena pubescens nigrocoerulea apicipennis caesus caesus variolosus beyeri gloriosa lecontei resplendens woodi agenor bottimeri femorata woodfordi hebraea maculatissima rusticola aequinoctialis balteata scalaris sp. 1 alba arguta autumnalis cavifrons comes crinita decimlineata devestiva diffracta fullo fullo gracilis hammondi nigra occidentalis sobrina tridentata variolosa punctata atrocoerulea atrocoerulea bipunctata cyanea formosana histeroidea indigonacea japonica mongolica Blackburn Wiedemeyer Wiedemeyer Lewis Burmeister Afzeli Cartwright Cartwright Cartwright Bates Cartwright Chapin Horn Waterhouse Reitter Creutzer Creutzer Kolenati Skinner LeConte Horn Boucard Horn Olivier Howden Waterhouse Waterhouse Olivier Bohmann Burmeister Olivier DeGeer Gory and Percheron Young Reitter Casey Schaeffer LeConte Casey LeConte Say Young Casey Linnaeus Linnaeus Horn LeConte Casey Linnaeus Casey Reitter Hentz Donovan Bates Kraatz Fabricius Hope Arrow Gyllenhal Motschulsky Newman Arrow 1885 1821 1821 1879 1844 1817 1948 1948 1948 1887 1948 1867 1841 1897 1796 1896 1846 1905 1854 1882 1875 1883 1789 1958 1894 1887 1789 1860 1842 1789 1778 1833 1 1 4 2 2 1 3 2 35 5 2 2 1 2 2 2 31 1 156 223 149 1 27 7 1 1 1 4 1 1 3 1 1 1 1 1 1 8 1 7 7 58 1 78 1903 1914 1927 1854 1914 1856 1824 1966 1891 1728 1728 15 1881 4 1856 172 1914 5 1767 8 1914 4 1890 1 1830 125 1805 2 1888 17 1829 1 1781 2 1831 11 1913 11 1817 7 1853 11 1838 272 1913 2 aus afr afr afr afr afr nea nea nea nea nea nea nea afr pal pal nea pal nea nea nea neo nea neo nea aus aus afr afr ori afr afr pal afr nea pal nea nea nea nea nea nea nea nea nea pal nea nea nea nea nea pal nea aus ori pal afr ori ori ori ori nea ori Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Popillia Popillia Popillia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Protaetia Psammodius Psammodius Pseudadoretus Pseudataenius Pseudinca Pseudocanthon Pseudochalcothea Pseudocotalpa Pseudocotalpa Pseudoprotaetia mutans pustulata quadriguttata acuminata aerata aeruginosa aeruginosa affinis affinis afflicta angustata angustata anovittata bakeri brevitaris brevitarsis catharica culta cuprea cuprea cuprea cuprea cuprea cuprea cuprea cuprea cuprea cuprea ducalis ferruginea funebris funesta fusca hungarica hungarica hungarica incerta judith karelini lugubris lugubris morio oblonga obscura opaca papuana phillippensis sp. sp. sp. speciosa speculifera vidua asper spp. phthisicus socialis admixtus perplexus pomacea andrewsi giulianii stolata Newman Fairmaire Fabricius Fabricius Erichson Drury Scopoli Andersch Reitter Gory and Percheron Germar Reitter Chevrolat Moser Kraatz Lewis Bates Waterhouse Fabricius Andersch Herbst Gory and Percheron Ball Paykull Andersch Andersch Reitter Falderman Mohnike Gory and Percheron Gory and Percheron Menetries Herbst Herbst Menetries Reitter Mohnike Reiche Zoubkoff Herbst Fabricius Fabricius Gory and Percheron Andersch Fabricius Moser Fabricius 1838 1887 1787 1775 1834 1770 1786 1797 1891 1833 1817 1902 1841 1917 1883 1879 1890 1879 1775 1797 1790 1833 1870 1799 1797 1797 1898 1835 1873 1833 1833 1836 1790 1790 1832 1901 1873 1871 1829 1786 1792 1781 1833 1797 1781 1914 1775 Gory and Percheron Swartz Waltl Fabricius 1833 1817 1838 1775 Dohrn Horn Hope LeConte Bates Hardy Hardy Olivier 1882 1871 1842 1847 1889 1971 1974 1789 16 14 8 4 1 5 1 4 1 3 6 1 1 1 1 22 1 2 7 4 8 3 1 13 1 6 1 1 1 7 1 2 97 13 1 1 1 1 2 1 3 5 2 1 3 5 7 1 2 5 1 4 4 6 10 1 9 1 38 1 1 2 8 ori ori ori ori ori ori pal pal pal pal pal pal ori ori ori afr ori ori pal pal pal pal pal pal pal pal pal pal ori ori pal pal ori pal afr afr ori pal afr ori pal pal pal pal pal ori ori ori pal ori ori ori pal pal pal pal nea afr nea ori nea nea afr Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Pseudotrematodes Ptenomela Pucaya Pycnoschema Pyronota Reineria Reineria Rhabdotis Rhizoplatys Rhizotrogus Rhizotrogus Rhizotrogus Rhizotrogus Rhizotrogus Rhizotrogus Rhomborhina Rhomborhina Rhomborhina Rhyparus Rhyssemodes Rhyssemus Rhyssemus Rhyssemus Rhyssemus Rhyssemus Rhyssemus Rutela Rutela Rutela Rutela Saprosites Scapanes Scaptobius Scarabaeus Scarabaeus Scarabaeus Scarabaeus Scarabaeus Scarabaeus Scarabaeus Scarabaeus Scarabaeus Scarabaeus Scatimus Sclerostomus Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica frivaldszkyi gratiosa pulchra sp. festiva margaritacea tigrina semipunctata mucronatus aequinoctialis aestivus maculicollis marginipes tusculus vernus japonica sp. unicolor risbeci orientalis californicus germanus puncticollis scaber sonatus spp. dorcyi formosa laeta lineola sp. australis pentarthrius cicatricosus fritschi infernalis intricatus laticollis pius puncticollis sacer semipunctatus variolosus ovatus costatus acicula alternata alternata alternata anthracina apatela aspera atracapilla brunnea campestris carolina concinna curvata diablo elusa falli fimbriata floridana Menetries Sharp Arrow 1836 1877 1911 Fabricius Mohnike Mohnike Fabricius Beauvois Herbst Olivier Villa Perris Buquet Germar Hope 1873 1873 1787 1805 1790 1789 1833 1870 1840 1823 1841 Motschulsky Paulian,??? Mulsant and Godart Horn Linnaeus Brown Haldeman LeConte 1861 Olivier Burmeister Weber Linnaeus 1789 1844 1801 1767 Boisduval Westwood Lucas Harold Klug Fabricius Linnaeus Illiger Latreille Linnaeus Fabricius Fabricius Harold H. and W. Dawson LeConte Dawson Dawson LeConte Dawson Dawson Kirby Linnaeus Dawson Dawson Dawson LeConte Dawson Dawson Dawson LeConte Dawson 1832 1874 1846 1868 1855 1801 1767 1803 1819 1758 1792 1787 1862 1874 1871 1767 1929 1848 1881 8 4 1 1 5 4 1 7 1 6 11 1 1 1 4 1 5 5 20 2 3 3 3 5 1 2 1 1 278 78 1 9 1 1 1 1 2 2 4 5 13 8 9 pal neo neo afr aus ori ori afr afr pal pal pal pal pal pal ori ori ori aus pal nea pal nea nea nea neo neo neo neo neo aus afr pal afr afr afr pal pal pal pal pal pal 1 neo 1932 1 nea 1856 85 nea 1933 1 nea 1933 2 nea 1856 21 nea 1922 1 nea 1922 116 nea 1837 44 nea 1758 18 pal 1919 2 nea 1920 3 nea 1947 3 nea 1856 3 nea 1967 3 nea 1919 7 nea 1932 2 nea 1856 10 nea 1967 1 nea Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Serica Sericospilus Sisyphus Sisyphus Sisyraphora Sphenognathus Spilophorus Spilophorus Spilophorus Spodistes Stalagmosoma Stenocrates Stenocrates Stenocrates Stephanorrhina Stephanucha Stephanucha Stephanucha Strategus Strategus Strategus Strategus Strategus Strategus Strategus Strategus Strategus Strategus Strategus Strategus Strategus Strategus frosti frosti frosti georgiana georigana holosericea intermixta iricolor ligulata mixta opposita parallela peregrina perigonis porcula prava pusilla pusilla satrapa sculptilus seratina sericea similis solita sponsa spp. tristis trociformis trociformis trogiformis ventura vespertina advena quadricollis schaefferi tomentosa bellicosus lugubris plagosus sp. mniszechi albella laborator sp. spp. guttata areata pilipennis thoracica aleous aloeus aloeus aloeus antaeus antaeus centaurus cessus mormon splendens spp. spp. spp. validus Dawson Dawson Dawson Leng Dawson Scopoli Blatchley Say Dawson LeConte Dawson Casey Chapin Dawson Casey Dawson Dawson Dawson Dawson Dawson LeConte Illiger LeConte Dawson Dawson LeConte Burmeister Dawson Uhler Dawson Gyllenhal Sharp Gory Linnaeus Gory and Percheron Boileau Fabricius Bohmann Thomas Pallas Fabricius Olivier Fabricius Kraatz Casey Linnaeus Linnaeus Linnaeus Linnaeus Drury Drury Kolbe LeConte Burmeister Beauvois Fabricius 1967 1 nea 1967 18 nea 1967 26 nea 1911 45 nea 1921 94 nea 1772 6 pal 1910 46 nea 1824 57 nea 1932 4 nea 1856 2 nea 1921 9 nea 1884 27 nea 1938 27 nea 1920 2 nea 1884 135 nea 1933 1 nea 1922 2 nea 1922 8 nea 1947 4 nea 1922 4 nea 1856 3 nea 1802 85 nea 1895 1 nea 1922 4 nea 1919 6 nea 53 nea 1850 67 nea 1844 33 nea 1932 18 nea 1855 10 nea 1932 1 nea 1817 47 nea 4 aus 1833 2 afr 1758 5 pal 1833 4 afr 1 neo 1775 2 afr 1857 1 afr 1 afr 1860 1 neo 1781 1 afr 1775 28 neo 16 neo 10 neo 1789 25 afr 1775 6 nea 1883 3 nea 1915 5 nea 11 neo 1758 53 nea 1758 39 nea 1758 87 neo 1773 4 nea 1773 113 nea 1906 4 neo 1866 35 nea 1847 1 nea 1809 27 nea 6 38 nea 66 neo 1775 4 neo Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Strigidia Strigoderma Strigoderma Strigoderma Strigoderma Strigoderma Strigoderma Strigoderma Strigoderma Strigoderma Syrictes Tanyproctus Taurhina Taurocopris Taurocopris Telaugis Tephraea Tesarius Tesarius Tesarius Thronistes Thronistes Thyce Thyridium Tmesorrhina Torynorrhina Trichillum Trichiopsammobius Trichiorhyssemus Trichius Trichius Trichius Trichius Trichius Trichius Trichius Trichius Trichius Trichius Trichius Trichius Trichius Trichius Trichius Trichius Trichius Trichogomphus Trichogomphus Tricholepis Trichostetha Trigonopeltastes Trigonopeltastes Trigonopeltastes Trigonopeltastes Trigonophorus Trigonophorus Trigonophorus Trigonophorus Triodonta Triodonta Triodonta Trioplus Trissodon liturella arboricola protea pygmaea pygmaea pygmaea sallaei sulcipennis texana viridicollis syrichthus reichei longiceps cadmus mimas aenescens dichroa caelatus mcclayi oregonensis rouxi spp. squamicollis semicinctum iris hyacinthina pilosus brasiliensis riparius abdominalis affinis assimilis bibens fasciatus fasciatus lunulatus piger rufobrunneus sexualis sp. spp. succinctus texanus viridans zonatus zonatus excavatus mongol lactea signatus delta floridana sallaei sinaloensis dilutus gracilipes nepalensis varians cinctipennis morio nitidula cylindricus glaber Kirby Fabricius Burmeister Fabricius Ohaus Brown Bates Burmeister Casey Schaeffer Fabricius Rambur Kolbe Harold Linnaeus Burmeister Schaum LeConte Cartwright Cartwright Burmeister 1792 1844 1798 1905 1925 1888 1844 1915 1907 1775 1843 1892 1868 1781 1844 1844 1857 1955 1955 1847 LeConte Bates Fabricius Hope Robinson Petrovitz Horn Menetries Gory and Percheron Kirby Fabricius Linnaeus Mulsant Fabricius Fabricius Casey Bedel 1856 1888 1781 1841 Pallas Horn Kirby Germar Heer Mohnike Arrow Gory Fabricius Forster Casey Bates Howden Bourgoin Westwood Hope Bourgoin Lucas Fabricius Rossi Mannerheim Arrow 1781 1876 1837 1794 1841 1874 1908 1963 1871 1832 1833 1837 1775 1758 1842 1775 1775 1914 1906 1771 1909 1889 1968 1914 1845 1831 1914 1849 1798 1790 1829 1 55 21 69 2 2 3 20 2 3 1 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Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Scarabaeidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Trogidae Tropinota Tropinota Tropinota Tropinota Valgus Valgus Valgus Valgus Xyloryctes Xyloryctes Xyloryctes Xylotrupes Omorgus Omorgus Omorgus Omorgus Omorgus Omorgus Omorgus Omorgus Omorgus Omorgus Omorgus Omorgus Omorgus Polynoncus Polynoncus Polynoncus Polynoncus Polynoncus Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox Trox undetermined hirta hirta squalida vestita californicus canaliculatus hemipterus sp. jamaicensis spp. telephus gideon asper inflatus monachus nodosus persuberosus punctatus punctatus scabrosus scutellaris suberosus tesselatus tytus umbonatus aricensis brasiliensis bullatus chilensis pilularius aequalis atrox cadaverinus capillaris fabricii foveicollis foveicollis gemmulatus hamatus hispidus insularis laticollis plicatus sabulosus scaber sonorae sordidus spinulosus striatus sulcatus terrestris tuberculatus unistriatus variolatus spp. Poda Linnaeus Scopoli Say Horn Olivier Linnaeus 1761 1766 1783 1825 1870 1789 1758 Drury 1773 Burmeister Linnaeus LeConte Loomis Herbst Robinson Vaurie Germar LeConte Beauvois Say Fabricius LeConte Robinson LeConte Gutierrez Vaurie Curtis Harold Germar Say LeConte Illiger Say Reiche Harold Robinson Horn Robinson Pontoppidan Chevrolat LeConte Robinson Linnaeus Linnaeus LeConte LeConte Robinson Melsheimer Thunberg Say Degeer Beauvois Melsheimer 1847 1767 1854 1922 1790 1940 1962 1824 1854 1805 1823 1775 1854 1941 1854 1950 1962 1845 1872 1824 1831 1854 1801 1823 1853 1872 1940 1874 1940 1763 1864 1854 1940 1758 1767 1854 1854 1940 1846 1787 1825 1774 1805 1847 45 3 26 12 6 29 39 11 149 64 8 142 8 4 22 6 16 28 7 19 8 191 4 6 1 1 2 1 1 2 9 4 1 35 2 11 4 5 8 5 1 9 54 3 3 116 29 26 3 2 24 15 70 24 6 pal pal pal pal nea nea pal nea nea neo neo ori nea nea nea nea pal nea nea nea nea nea nea nea nea pal pal pal pal pal nea nea neo nea neo nea nea nea nea neo pal nea nea neo nea nea nea nea afr nea nea nea nea
Podobné dokumenty
COMMON NAME Coleoptera Carabidae 1 Omophron Coleoptera
Bembidion
Bembidion
Paratachys
Lebia
Axinopalpus
Cicindela
Peltodytes
Laccophilus
Stictotarsus
[Bidessini]
Ochthebius
Acrotrichis
Thinopinus
Hadrotes
Heterothops
Cafius
Cafius
Cafius
Bledius
Blediu...
Ants
(Hymenoptera)